But within the Despicable Me universe, the Minions are — or were — the product of Dr. Nefario’s genetic engineering efforts to manufacture an army of loyal workers for his villain-in-chief.

However, he later betrays him. El Macho is the villain of Despicable Me 2. In the film, he joins forces with Dr. Nefario in order to kidnap and mutate Gru’s minions with the help of the PX-41 ...